The domain within your query sequence starts at position 317 and ends at position 427; the E-value for the 4dex domain shown below is 5e-45.

EFAKERERVENRRAFLKLRRQQQIERELNGYMEWISKAEEVILAEDETDVEQRHPFDGAL
RRATLKKSKTDLLNPEEAEDQLADIASVGSPFARASIKSAKLENSTFFHKK

This is a PDB domain - please see the PDBSUM entry of 4dex for further annotation